top of page




Pimiento C, Albouy C, Silvestro D, Mouton TL, Velez L, Mouillot D, Griffin JN, Leprieur F.  2023. Functional diversity of sharks and rays is highly vulnerable and supported by unique species and locations worldwide. Nature Communications 14, 691 


Munoz, F., Klausmeier, C.A., Gaüzère, P., Kandlikar, G., Litchman, E., Mouquet, N. Ostling A, Thuiller W., Algar A, Auber A,. Cadotte M. Delalandre L., Denelle P.,  Enquist B.J.,  Fortunel C.,  Grenié M.,  Loiseau, N.,  Mahaut L.,  Maire M.,  Mouillot D.,  Pimiento C.,  Violle C.,  Kraft NJB.(2023) The ecological causes of functional distinctiveness in communities. Ecology Letters, 00, 1– 14. 

Villafaña JA, Rivadeneira MM, Pimiento C, Kriwet J. (2023) Diversification trajectories and paleobiogeography of Neogene chondrichthyans from Europe. Paleobiology 1-13. 


Cooper, J. A.,  Griffin, J. N.,  Kindlimann, R., &  Pimiento, C. (2023).  Are shark teeth proxies for functional traits? A framework to infer ecology from the fossil record. Journal of Fish Biology  1– 17. (Student-led paper).



Dillon E, Pier, P, Smith JA, Raja NB, Austin EL, Cybulski, JD, De Entrambasaguas J, Durham SR, Grether C, Haldar H, Kocáková, K, Lin CH, Mazzini I, Mychajliw A, Ollendorf, A, Pimiento C, Regalado-Fernandez OR, Smith, IE, Dietl GP (2022) What is conservation paleobiology? Tracking 20 years of research and development. Frontiers in Ecology and Evolution 10:1031483 https://doi:10.3389/fevo.2022.1031483

Pimiento, C & Antonelli, A (2022) Integrating deep-time palaeontology in conservation prioritisation. Frontiers in Ecology and Evolution 10:959364  


Joos, J, Pimiento C, Müller J, Miles D (2022) Quaternary megafauna extinctions altered body size distribution in tortoises. Proceedings of the Royal Society B 20221947 


Cooper JA, Hutchinson JR, Bernvi DC, Cliff, G, Wilson RW, Dicken ML, Menzel, J, Wroe, S, Pirlo, J and Pimiento, C. (2022) The extinct shark Otodus megalodon was a transoceanic super-predator: inferences from 3D modelling. Science Advances abm9424, 




Bazzi M, Campione N, Kear BP, Pimiento C, Ahlberg, PE (2021) Feeding ecology has shaped the evolution of modern sharks. Current Biology 31, 1-11 


Pimiento C. & Pyenson N. (2021). Perspective: When sharks nearly disappeared. Science 372, 1036-1037  


Paillard, A.,  Shimada, K.,  Pimiento, C.  (2020) The fossil record of extant elasmobranchs. Journal of   Fish Biology 1– 11. (Student-led paper).



Cooper, J.A., Pimiento, C., Ferrón, H.G.. Benton, M. (2020) Body dimensions of the extinct giant shark Otodus megalodon: a 2D reconstruction. Scientific Reports 10, 14596.  (Student-led paper).

Pimiento C, Bacon CD, Silvestro D, Hendy A, Jaramillo C, Zizka A, Meyer X, Antonelli A. (2020) Selective extinction against redundant species buffers functional diversity. Proceedings of the Royal Society B 287: 20201162.

Villafaña JA, Hernandez S, Alvarado A, Shimada K, Pimiento C, Rivadeneira MM, Kriwet J. (2020) First evidence of a palaeo-nursery area of the great white shark. Scientific Reports 10, 8502

Pimiento C, Leprieur F, Silvestro D, Lefcheck J, Albouy C, Rasher DB, Davis M, Svenning JC, Griffin JN. (2020) Functional diversity and the fate of marine megafauna in the Anthropocene. Science Advances 6, eaay7650 doi: 10.1126/sciadv.aay7650

Pimiento C, Benton M. (2020) The impact of the Pull of the Recent in extant elasmobranchs. Palaeontology 2020, pp. 1–6 DOI: 10.1111/pala.12478.



Pimiento C, Cantalapiedra LJ, Shimada K, Field, D, Smaers, J. (2019) Evolutionary pathways toward gigantism in sharks and rays. Evolution 73: 588-599 DOI: 10.1111/evo.13680.

Pimiento C. (2018) Perspective: Our shallow-water origins. Science 362 (6413): 8-9 DOI: 10.1126/science.aav3494.

Pimiento C, Tang KL, Zamora S, Klug C, Sánchez-Villagra MR. (2018) Assessing canalisation of intraspecific variation on a macroevolutionary scale: the case of crinoid arms through the Phanerozoic. PeerJ 6:e4899 DOI: 10.7717/peerj.4899.

Pimiento C, Griffin JN, Clements CF, Silvestro D, Varela S, Uhen M, Jaramillo C (2017) The Pliocene marine megafauna extinction and its impact on functional diversity. Nature Ecology and Evolution 1(8) 1100.  DOI: 10.1038/s41559-017-0223-6 (Full text).

Perez, V., Pimiento, C., Hendy, A., González-Barba, G., Hubbell, G., & MacFadden, B. (2017). Late Miocene chondrichthyans from Lago Bayano, Panama: Functional diversity, environment and biogeography. Journal of Paleontology, 1-36.  DOI: 10.1017/jpa.2017.5. 


Pimiento C, MacFadden B, Clements CF, Varela S, Jaramillo C, Velez-Juarbe J, Silliman BR (2016) Geographical distribution patterns of Carcharocles megalodon over time reveal clues about extinction mechanisms. Journal of Biogeography 43(8): 1645–1655. DOI: 10.1111/jbi.12754.

Velez-Juarbe J, Wood AR., Pimiento C. (2016) Pygmy sperm whales (Odontoceti, Physeteroidea, Kogiidae) from the early Pliocene of Florida. Journal of Vertebrate Paleontology 36(4): e1135806 DOI: 10.1080/02724634.2016.1135806.

Pimiento C, Nifong JC, Hunter ME, Monaco E, Silliman BR (2015) Habitat use patterns of the invasive red lionfish Pterois volitans: a comparison between mangrove and reef systems in San Salvador, Bahamas. Marine Ecology 36:28–37 DOI: 10.1111/maec.12114.

MacFadden BJ, Symister C, Cannarozzi N, Pimiento C, De Gracia C. (2015) Comparative diagenesis and rare earth element (REE) variation in Miocene invertebrate and vertebrate fossils from Panama. Journal of Geology 123(6):491-507 DOI: 10.1086/684006.

Pimiento C & Balk MA (2015) Body size trends of the extinct giant shark Carcharocles megalodon: A deep-time perspective on marine apex predators. Paleobiology (41):479-490 DOI: 10.1017/pab.2015.16.

Carrillo-Briceño JD, De Gracia C, Pimiento C, Aguilera OA, Kindlimann R, Santamaría P, Jaramillo C (2015) New Late Miocene Chondrichthyan Assemblage from the Chagres Formation, Panama. Journal of South American Earth Sciences 60:56–70. DOI: 10.1016/j.jsames.2015.02.001.



Pimiento C & Clements C (2014) When did Carcharocles megalodon become extinct? A new analysis of the fossil record. PLoSONE 9(10):e111086  DOI: 10.1371/journal.pone.0111086.

Vasquez S & †Pimiento C (2014) Sharks and Rays from the Tonosi Formation (Eocene of Panama). The Central American Journal of Geology. 51:165–169 DOI: 10.15517/rgac.v51i1.16911.

Pimiento C, Gonzalez-Barba G, Ehret DJ, MacFadden BJ, Hendy A, Jaramillo C (2013) Sharks and Rays (Chondrichthyes, Elasmobranchii) from the late Miocene Gatun Formation of Panama. Journal of Paleontology 87(5):55–774. DOI: 10.1666/12-117.

Pimiento C, Gonzalez G, Hendy A, Jaramillo C, MacFadden BJ, Montes C, Suarez SC, Shippritt M (2013) Early Miocene chondrichthyans from the Culebra Formation, Panama: A window into marine vertebrate faunas before closure of the Central American Seaway. Journal of South American Earth Sciences 42:159-170. DOI: 10.1016/j.jsames.2012.11.005.

Pimiento C, Monaco E, Barbour A, Silliman BR (2013) Predation on speared lionfish Pterois volitans by spotted moray Gymnothorax moringa. Florida Scientist 76(3-4):391–394.

Pimiento C, Ehret DJ, MacFadden BJ, Hubbell G (2010) Ancient Nursery Area for the Extinct Giant Shark Megalodon from the Miocene of Panama. PLoSONE 5(5):e10552 DOI: 10.1371/journal.pone.0010552.

Uhen MD, Coates AG, Jaramillo CA, Montes C, Pimiento C, Rincon A, Strong N, Velez-Juarbe J (2010) Marine mammals from the Miocene of Panama. Journal of South American Earth Sciences 30 (3-4):167–175 DOI:10.1016/j.jsames.2010.08.002.

In Education and Outreach

Pimiento C (2015) Engaging students in paleontology: The design and implementation of an undergraduate-level blended course in Panama. Evolution: Education and Outreach 8:19 DOI:

Pimiento C & Pringle R (2011) Fossil sharks: Learning from and about the past. Science Scope 34(6): 2–59.


Griffin JN, Leprieur F, Silvestro D, Lefcheck JS, Albouy C, Rasher DB, Davis M, Svenning J-C, Pimiento C (2020) Functionally unique, specialised, and endangered (FUSE) species: towards integrated metrics for the conservation prioritisation toolbox. bioRxiv 2020.05.09.084871; doi: 

bottom of page